RESULTS OF LARGE MAMMAL MONITORING IN THE ROZTOCHYA NATURE RESERVE OVER 40 YEARS OF OBSERVATIONS, 1984–2025
DOI: http://dx.doi.org/10.30970/sbi.2001.865
Abstract
Background. Long-term wildlife monitoring is crucial for understanding population dynamics and informing conservation strategies. This study presents a 40-year analysis of large mammal communities in the Roztochya Nature Reserve (Western Ukraine). The study period covers two distinct methodological approaches: traditional surveys (1984–2019), which yielded data with significant limitations for quantitative analysis, and modern camera trap monitoring initiated in 2020 to overcome these shortcomings and provide objective, verifiable data on the region’s fauna.
Materials and Methods. Historical data from 1984–2019, collected via transect counts and track surveys, were critically reviewed. From 2020 to mid-2025, a systematic camera trap survey was implemented, with the network expanding from 4 to 20 devices. Analysis focused on species composition, detection frequencies, and seasonal activity patterns. We calculated a relative abundance index to assess recent population trends and compared the effectiveness of the two monitoring approaches.
Results. The camera trap monitoring yielded over 2000 detection days for 13 mammal species. The European roe deer (Capreolus capreolus) was the most frequent species (~53.4 %), followed by the red fox (Vulpes vulpes, ~18 %) and the wild boar (Sus scrofa, ~10 %). A significant increase in detections was observed for roe deer and wild boar in 2023–2024. A key finding was the confirmation of the European wildcat’s (Felis silvestris) regular presence since 2024, changing its status from extremely rare to a permanent resident. The transient status of the grey wolf (Canis lupus) was also confirmed.
Conclusions. The transition to camera trapping provides a more accurate assessment of the mammal community, proving highly effective for detecting rare and elusive species. These results allow for a critical revision of the conservation status of several key species, most notably the European wildcat. The study underscores the importance of the Roztochya Nature Reserve as a key refugium and ecological corridor, with population dynamics likely influenced by recent nationwide changes in hunting management.
Keywords
Full Text:
PDFReferences
| Akimov, I. A. (Ed.). (2009). Chervona knyha Ukrainy. Tvarynnyi svit [Red Data Book of Ukraine. The animal world]. Kyiv: Globalkonsalting. Google Scholar | ||||
| ||||
| Albon, S. D., & Langvatn, R. (1992). Plant phenology and the benefits of migration in a temperate ungulate. Oikos, 65(3), 502-513. doi:10.2307/3545568 Crossref ● Google Scholar | ||||
| ||||
| Andriienko, T. L., Popovych, S. Yu., Parchuk, H. V., Havrylenko, V. S., & Priadko, O. I. (2002). Prohrama Litopysu pryrody dlia zapovidnykiv ta natsionalnykh pryrodnykh parkiv [Program of the Chronicle of Nature for Nature Reserves and National Nature Parks]. Kyiv: Akademperiodyka. (In Ukrainian) Google Scholar | ||||
| ||||
| Apollonio, M., Andersen, R., & Putman, R. (Eds.). (2010). European ungulates and their management in the 21st century. Cambridge University Press. Google Scholar | ||||
| ||||
| Baker, P. J., Funk, S. M., Harris, S., & White, P. C. (2000). Flexible spatial organization of urban foxes, Vulpes vulpes, before and during an outbreak of sarcoptic mange. Animal Behaviour, 59(1), 127-146. doi:10.1006/anbe.1999.1285 Crossref ● PubMed ● Google Scholar | ||||
| ||||
| Baubet, E., Bonenfant, C., & Brandt, S. (2004). Diet of the wild boar in the French Alps. Galemys, 16(1), 101-113. doi:10.7325/galemys.2004.ne.a8 Crossref ● Google Scholar | ||||
| ||||
| Best, D. A., Lynch, G. M., & Rongstad, O. J. (1978). Seasonal activity patterns of moose in the Swan Hills, Alberta. Alces, 14, 109-125. Google Scholar | ||||
| ||||
| Bieber, C., & Ruf, T. (2005). Population dynamics in wild boar Sus scrofa: ecology, elasticity of growth rate and implications for the management of pulsed resource consumers. Journal of Applied Ecology, 42(6), 1203-1213. doi:10.1111/j.1365-2664.2005.01094.x Crossref ● Google Scholar | ||||
| ||||
| Bobek, B., Furtek, J., Merta, D., & Wojciuch-Płoskonka, M. (2024). The influence of forest site types on the distribution of moose Alces alces in north-eastern Poland. Scandinavian Journal of Forest Research, 39(5), 257-262. doi:10.1080/02827581.2024.2388113 Crossref ● Google Scholar | ||||
| ||||
| Bonenfant, C., Loe, L. E., Mysterud, A., Langvatn, R., Stenseth, N. C., Gaillard, J. M., & Klein, F. (2004). Multiple causes of sexual segregation in European red deer: enlightenments from varying breeding phenology at high and low latitude. Proceedings of the Royal Society of London. Series B: Biological Sciences, 271(1542), 883-892. doi:10.1098/rspb.2003.2661 Crossref ● PubMed ● PMC ● Google Scholar | ||||
| ||||
| Boreiko, V. Ye., & Parnikhoza, I. Yu. (2023, August 2). Pidstavy dlia okhorony losia v Ukraini [Rationale for protection of the moose in Ukraine]. Expediciya XXI. Retrieved from https://expedicia.org/pidstavi-dlya-okhoroni-losya-v-ukraini | ||||
| ||||
| Bowyer, R. T. (2022). Sexual segregation in ungulates: ecology, behavior, and conservation. Baltimore: Johns Hopkins University Press. doi:10.56021/9781421445076 Crossref ● Google Scholar | ||||
| ||||
| Broders, H. G., Coombs, A. B., & McCarron, J. R. (2012). Ecothermic responses of moose (Alces alces) to thermoregulatory stress on mainland Nova Scotia. Alces, 48, 53-61. Google Scholar | ||||
| ||||
| Cano-Martínez, R., Carricondo-Sanchez, D., Devineau, O., & Odden, M. (2021). Small rodent cycles influence interactions among predators in a boreal forest ecosystem. Mammal Research, 66(4), 583-593. doi:10.1007/s13364-021-00590-7 Crossref ● Google Scholar | ||||
| ||||
| Carbone, C., Christie, S., Conforti, K., Coulson, T., Franklin, N., Ginsberg, J. R., ... & Wan Shahruddin, W. N. (2001). The use of photographic rates to estimate densities of tigers and other cryptic mammals. Animal Conservation, 4(1), 75-79. doi:10.1017/s1367943001001081 Crossref ● Google Scholar | ||||
| ||||
| Castañeda, I., Doherty, T. S., Fleming, P. A., Stobo-Wilson, A. M., Woinarski, J. C., & Newsome, T. M. (2022). Variation in red fox Vulpes vulpes diet in five continents. Mammal Review, 52(3), 328-342. doi:10.1111/mam.12292 Crossref ● Google Scholar | ||||
| ||||
| Cavallini, P., & Lovari, S. (1991). Environmental factors influencing the use of habitat in the red fox, Vulpes vulpes. Journal of Zoology, 223(2), 323-339. doi:10.1111/j.1469-7998.1991.tb04768.x Crossref ● Google Scholar | ||||
| ||||
| Cavallini, P., & Santini, S. (1995). Timing of reproduction in the Red fox, Vulpes vulpes. Zeitschrift fur Saugetierkunde, 60(6), 337-342. Google Scholar | ||||
| ||||
| Chapron, G., Kaczensky, P., Linnell, J. D., von Arx, M., Huber, D., Andrén, H., ... & Boitani, L. (2014). Recovery of large carnivores in Europe's modern human-dominated landscapes. Science, 346(6216), 1517-1519. doi:10.1126/science.1257553 Crossref ● PubMed ● Google Scholar | ||||
| ||||
| Clutton-Brock, T. H., Guinness, F. E., & Albon, S. D. (1982). Red deer: behavior and ecology of two sexes. University of Chicago Press. Google Scholar | ||||
| ||||
| Convention on the Conservation of European Wildlife and Natural Habitats. (1979, accessed 21 March 2025). Annex II - strictly protected fauna species. Retrieved from https://rm.coe.int/168078e2ff | ||||
| ||||
| Debeffe, L., Focardi, S., Bonenfant, C., Hewison, A. J., Morellet, N., Vanpé, C., ... & Cagnacci, F. (2014). A one night stand? Reproductive excursions of female roe deer as a breeding dispersal tactic. Oecologia, 176(2), 431-443. doi:10.1007/s00442-014-3021-8 Crossref ● PubMed ● Google Scholar | ||||
| ||||
| Debeffe, L., Maren Rivrud, I., Brekkum, Ø., Meisingset, E. L., & Mysterud, A. (2017). Implications of the forage maturation hypothesis for activity of partially migratory male and female deer. Ecosphere, 8(12), e02050. doi:10.1002/ecs2.2050 Crossref ● Google Scholar | ||||
| ||||
| Doherty, T. S., Dickman, C. R., Glen, A. S., Newsome, T. M., Nimmo, D. G., Ritchie, E. G., ... & Wirsing, A. J. (2017). The global impacts of domestic dogs on threatened vertebrates. Biological Conservation, 210, 56-59. doi:10.1016/j.biocon.2017.04.007 Crossref ● Google Scholar | ||||
| ||||
| Dolapchiev, N., Popova, E., & Chanev, M. (2024). Spatial and temporal relationships between predators in the "Sinite Kamani" Nature Park, Bulgaria. Ecologia Balkanica, 16(2), 70-79. Google Scholar | ||||
| ||||
| Dykyi, I. V. (2001). Current state of the badger (Meles meles L.) population in Western Ukraine. Visnyk of Lviv University. Biological series, 27, 151-155. (In Ukrainian) Google Scholar | ||||
| ||||
| Dziki-Michalska, K., Tajchman, K., & Budzynska, M. (2019). Increase in the moose (Alces alces L. 1758) population size in Poland: causes and consequences. Annals of Warsaw University of Life Sciences - SGGW. Animal Science, 58(3), 203-214. doi:10.22630/aas.2019.58.3.20 Crossref ● Google Scholar | ||||
| ||||
| European Food Safety Authority (EFSA), Ståhl, K., Boklund, A. E., Podgórski, T., Vergne, T., Abrahantes, J. С., & Mur, L. (2024). Epidemiological analysis of African swine fever in the European Union during 2023. EFSA Journal, 22(5), e8809. doi:10.2903/j.efsa.2025.9436 Crossref ● PubMed ● PMC ● Google Scholar | ||||
| ||||
| Franzmann, A. W. (1981). Alces alces. Mammalian Species, 154, 1-7. doi:10.2307/3503876 Crossref ● Google Scholar | ||||
| ||||
| Gamelon, M., Touzot, L., Baubet, É., Cachelou, J., Focardi, S., Franzetti, B., Nivois, É., Veylit, L., & Sæther, B. (2021). Effects of pulsed resources on the dynamics of seed consumer populations: a comparative demographic study in wild boar. Ecosphere, 12(5), e03395 doi:10.1002/ecs2.3395 Crossref ● Google Scholar | ||||
| ||||
| Goszczyński, J., Juszko, S., Pacia, A., & Skoczyńska, J. (2005). Activity of badgers (Meles meles) in Central Poland. Mammalian Biology, 70(1), 1-11. doi:10.1078/1616-5047-00171 Crossref ● Google Scholar | ||||
| ||||
| Huzii, A. I. (1997). Fauna i naselennia khrebetnykh Zakhidnoho rehionu Ukrainy. Tom 1. Roztochchia [Fauna and populations of vertebrates of the western region of Ukraine. Vol. 1. Roztochia]. Kyiv: Ukrainian State Forestry University. (In Ukraine) Google Scholar | ||||
| ||||
| Henttonen, H., Kranz, A., Stubbe, M., Maran, T., & Tikhonov, A. (2007). Alces alces ssp. alces (Europe assessment). The IUCN Red List of Threatened Species. Version 2007: e.T41782A10539156 Google Scholar | ||||
| ||||
| Johnson, D. D., Jetz, W., & Macdonald, D. W. (2002). Environmental correlates of badger social spacing across Europe. Journal of Biogeography, 29(3), 411-425. doi:10.1046/j.1365-2699.2002.00680.x Crossref ● Google Scholar | ||||
| ||||
| Kays, R., Arbogast, B. S., Baker-Whatton, M., Beirne, C., Boone, H. M., Bowler, M., ... & Spironello, W. R. (2020). An empirical evaluation of camera trap study design: how many, how long and when? Methods in Ecology and Evolution, 11(6), 700-713. doi:10.1111/2041-210x.13370 Crossref ● Google Scholar | ||||
| ||||
| Kelly, D. (1994). The evolutionary ecology of mast seeding. Trends in Ecology & Evolution, 9(12), 465-470. doi:10.1016/0169-5347(94)90310-7 Crossref ● PubMed ● Google Scholar | ||||
| ||||
| King, C. M., Powell, R. A., & Powell, C. (2006). The natural history of weasels and stoats: ecology, behavior, and management. New York: Oxford University Press. doi:10.1093/acprof:oso/9780195322712.001.0001 Crossref ● Google Scholar | ||||
| ||||
| Kobryn, H. T., Swinhoe, E. J., Bateman, P. W., Adams, P. J., Shephard, J. M., & Fleming, P. A. (2023). Foxes at your front door? Habitat selection and home range estimation of suburban red foxes (Vulpes vulpes). Urban Ecosystems, 26(1), 1-17. doi:10.1007/s11252-022-01252-5 Crossref ● Google Scholar | ||||
| ||||
| Kowalczyk, R., Jędrzejewska, B., & Zalewski, A. (2003). Annual and circadian activity patterns of badgers (Meles meles) in Białowieża Primeval Forest (eastern Poland) compared with other Palaearctic populations. Journal of Biogeography, 30(3), 463-472. doi:10.1046/j.1365-2699.2003.00804.x Crossref ● Google Scholar | ||||
| ||||
| Kropil, R., Smolko, P., & Garaj, P. (2015). Home range and migration patterns of male red deer Cervus elaphus in Western Carpathians. European Journal of Wildlife Research, 61(1), 63-72. doi:10.1007/s10344-014-0874-4 Crossref ● Google Scholar | ||||
| ||||
| Kruuk, H., & Parish, T. (1981). Feeding specialization of the European badger Meles meles in Scotland. The Journal of Animal Ecology, 50(3), 773-788. doi:10.2307/4136 Crossref ● Google Scholar | ||||
| ||||
| Linnell, J. D. C., & Andersen, R. (1998). Territorial fidelity and tenure in roe deer bucks. Acta Theriologica, 43, 67-75. doi:10.4098/at.arch.98-5 Crossref ● Google Scholar | ||||
| ||||
| Loe, L. E., Bonenfant, C., Mysterud, A., Severinsen, T., Oristland, N. A., Langvatn, R., & Stenseth, N. C. (2005). Climate predictability and breeding phenology in red deer: timing and synchrony of calving. Journal of Animal Ecology, 74(4), 579-588. doi:10.1111/j.1365-2656.2005.00987.x Crossref ● Google Scholar | ||||
| ||||
| Maran, T., Skumatov, D., Gomez, A., Põdra, M., Abramov, A. V., & Dinets, V. (2016). Mustela lutreola. The IUCN red list of threatened species 2016: E. T14018A45199861. Accessed on January, 5, 2024. doi:10.2305/iucn.uk.2016-1.rlts.t14018a45199861.en Crossref ● Google Scholar | ||||
| ||||
| Martsiv, M., Syrota, Y., & Dykyy, I. (2021). Diet composition of the red fox, Vulpes vulpes Linnaeus, 1758 (Canidae, Carnivora) in Western Ukraine. Acta Biologica Universitatis Daugavpiliensis, 21(1), 71-81. Google Scholar | ||||
| ||||
| Marynych, O. M. (Ed.). (1993). Heohrafichna entsyklopediia Ukrainy [Geographical encyclopedia of Ukraine] (Vols. 1-3). Kyiv: Ukrainian Soviet Encyclopedia named after M. P. Bazhan. (In Ukraine) Google Scholar | ||||
| ||||
| Mattucci, F., Oliveira, R., Lyons, L. A., Alves, P. C., & Randi, E. (2016). European wildcat populations are subdivided into five main biogeographic groups: consequences of Pleistocene climate changes or recent anthropogenic fragmentation? Ecology and Evolution, 6(1), 3-22. doi:10.1002/ece3.1815 Crossref ● PubMed ● PMC ● Google Scholar | ||||
| ||||
| McClune, D. W., Kostka, B., Delahay, R. J., Montgomery, W. I., Marks, N. J., & Scantlebury, D. M. (2015). Winter is coming: seasonal variation in resting metabolic rate of the European badger (Meles meles). PLoS One, 10(9), e0135920. doi:10.1371/journal.pone.0135920 Crossref ● PubMed ● PMC ● Google Scholar | ||||
| ||||
| McLeod, A. I. (2022). Kendall: Kendall rank correlation and Mann–Kendall trend test (Version 2.2.1). doi:10.32614/cran.package.kendall Crossref ● Google Scholar | ||||
| ||||
| Melis, C., Cagnacci, F., & Lovari, S. (2004). Site fidelity of male roe deer in a Mediterranean fragmented area. Hystrix, 15(1), 63-68. Google Scholar | ||||
| ||||
| Mikulka, O., Zeman, J., Drimaj, J., Plhal, R., Adamec, Z., Kamler, J., & Heroldová, M. (2018). The importance of natural food in wild boar (Sus scrofa) diet during autumn and winter. Folia Zoologica, 67(3-4), 165-172. doi:10.25225/fozo.v67.i3-4.a3.2018 Crossref ● Google Scholar | ||||
| ||||
| Ministry of Environmental Protection and Natural Resources of Ukraine. (2021, January 19). Pro zatverdzhennia perelikiv vydiv tvaryn, shcho zanosiatsia do Chervonoi knyhy Ukrainy (tvarynnyi svit), ta vydiv tvaryn, shcho vykliucheni z Chervonoi knyhy Ukrainy (tvarynnyi svit) [On approval of the lists of animal species included in the Red Data Book of Ukraine (Animal World), and those excluded from it (Animal World)] (Order No. 29). Retrieved from https://zakon.rada.gov.ua/laws/show/z0260-21#Text | ||||
| ||||
| Morelle, K., Podgórski, T., Prévot, C., Keuling, O., Lehaire, F., & Lejeune, P. (2015). Towards understanding wild boar Sus scrofa movement: a synthetic movement ecology approach. Mammal Review, 45(1), 15-29. doi:10.1111/mam.12028 Crossref ● Google Scholar | ||||
| ||||
| Mysterud, A., Loe, L. E., Zimmermann, B., Bischof, R., Veiberg, V., & Meisingset, E. (2011). Partial migration in expanding red deer populations at northern latitudes - a role for density dependence? Oikos, 120(12), 1817-1825. doi:10.1111/j.1600-0706.2011.19439.x Crossref ● Google Scholar | ||||
| ||||
| O'Connell, A. F., Nichols, J. D., & Karanth, K. U. (Eds.). (2011). Camera traps in animal ecology: methods and analyses (Vol. 271). New York: Springer. doi:10.1007/978-4-431-99495-4 Crossref ● Google Scholar | ||||
| ||||
| Oehler, F., Hagen, R., Hackländer, K., Walton, Z., Ashish, K., & Arnold, J. (2025). How do red foxes (Vulpes vulpes) explore their environment? Characteristics of movement patterns in time and space. Movement Ecology, 13(1), 4. doi:10.1186/s40462-024-00526-1 Crossref ● PubMed ● PMC ● Google Scholar | ||||
| ||||
| Pagon, N., Grignolio, S., Brivio, F., Marcon, A., & Apollonio, M. (2017). Territorial behaviour of male roe deer: a telemetry study of spatial behaviour and activity levels. Folia Zoologica, 66(4), 267-276. doi:10.25225/fozo.v66.i4.a9.2017 Crossref ● Google Scholar | ||||
| ||||
| Panzacchi, M., Linnell, J. D. C., Odden, M., Odden, J., & Andersen, R. (2009). Habitat and roe deer fawn vulnerability to red fox predation. Journal of Animal Ecology, 78(6), 1124-1133. doi:10.1111/j.1365-2656.2009.01584.x Crossref ● PubMed ● Google Scholar | ||||
| ||||
| Parker, K. L., Gillingham, M. P., Hanley, T. A., & Robbins, C. T. (1999). Energy and protein balance of free-ranging black-tailed deer in a natural forest environment. Wildlife Monographs, 143, 3-48. Google Scholar | ||||
| ||||
| Pearse, I. S., Koenig, W. D., & Kelly, D. (2016). Mechanisms of mast seeding: resources, weather, cues, and selection. New Phytologist, 212(3), 546-562. doi:10.1111/nph.14114 Crossref ● PubMed ● Google Scholar | ||||
| ||||
| Pejsak, Z., Truszczyński, M., Niemczuk, K., Kozak, E., & Markowska-Daniel, I. (2018). Epidemiology of African swine fever in Poland since the detection of the first case. Polish Journal of Veterinary Sciences, 21(1), 119-125. doi:10.2478/pjvs-2014-0097 Crossref ● PubMed ● Google Scholar | ||||
| ||||
| Plhal, R., Kamler, J., Homolka, M., & Adamec, Z. (2011). An assessment of the applicability of photo trapping to the wild boar population density in a forest environment. Folia Zoologica, 60(3), 237-246. doi:10.25225/fozo.v60.i3.a8.2011 Crossref ● Google Scholar | ||||
| ||||
| Pryrodnyi zapovidnyk "Roztochchya" [Roztochya Nature Reserve]. Litopys pryrody [Chronicle of nature]. (1986-2024). Annual reports submitted to the Ministry of Environmental Protection and Natural Resources of Ukraine. Ivano-Frankove: Roztochya Nature Reserve. (In Ukrainian) | ||||
| ||||
| R Core Team. (2023). R: a language and environment for statistical computing. R Foundation for Statistical Computing. Retrieved from https://www.R-project.org | ||||
| ||||
| Rigoudy, N., Dussert, G., Benyoub, A., Besnard, A., Birck, C., Boyer, J., ... & Chamaillé-Jammes, S. (2023). The DeepFaune initiative: a collaborative effort towards the automatic identification of European fauna in camera trap images. European Journal of Wildlife Research, 69(6), 113. do:10.1007/s10344-023-01742-7 Crossref ● Google Scholar | ||||
| ||||
| Rodríguez, A., Urra, F., Jubete, F., Román, J., Revilla, E., & Palomares, F. (2020). Spatial segregation between red foxes (Vulpes vulpes), European wildcats (Felis silvestris) and domestic cats (Felis catus) in pastures in a livestock area of Northern Spain. Diversity, 12(7), 268. doi:10.3390/d12070268 Crossref ● Google Scholar | ||||
| ||||
| Rovero, F., & Zimmermann, F. (Eds.). (2016). Camera trapping for wildlife research. Pelagic Publishing. doi:10.2307/jj.28833751 Crossref ● Google Scholar | ||||
| ||||
| Sauter-Louis, C., Conraths, F. J., Probst, C., Blohm, U., Schulz, K., Sehl, J., ... & Blome, S. (2021). African swine fever in wild boar in Europe - a review. Viruses, 13(9), 1717. doi:10.3390/v13091717 Crossref ● PubMed ● PMC ● Google Scholar | ||||
| ||||
| Servanty, S., Gaillard, J. M., Toïgo, C., Brandt, S., & Baubet, E. (2011). Pulsed resources and climate-induced variation in the reproductive traits of wild boar under high hunting pressure. Journal of Animal Ecology, 80(6), 1278-1290. doi:10.1111/j.1365-2656.2009.01579.x Crossref ● PubMed ● Google Scholar | ||||
| ||||
| Soininen, E. M., & Neby, M. (2024). Small rodent population cycles and plants - after 70 years, where do we go? Biological Reviews, 99(1), 265-294. doi:10.1111/brv.13021 Crossref ● PubMed ● Google Scholar | ||||
| ||||
| Stache, A., Heller, E., Hothorn, T., & Heurich, M. (2013). Activity patterns of European roe deer (Capreolus capreolus) are strongly influenced by individual behaviour. Folia Zoologica, 62(1), 67-75. doi:10.25225/fozo.v62.i1.a10.2013 Crossref ● Google Scholar | ||||
| ||||
| Sugianto, N. A., Newman, C., Macdonald, D. W., & Buesching, C. D. (2023). Effects of weather and social factors on hormone levels in the European badger (Meles meles). Zoology, 158, 126093. doi:10.1016/j.zool.2023.126093 Crossref ● Google Scholar | ||||
| ||||
| Tatarynov, K. A. (1973). Fauna khrebetnykh zakhodu Ukrainy [Vertebrate Fauna of Western Ukraine]. Lviv: Lviv University Press. (In Ukraine) Google Scholar | ||||
| ||||
| Thurfjell, H., Ball, J. P., Åhlén, P. A., Kornacher, P., Dettki, H., & Sjöberg, K. (2009). Habitat use and spatial patterns of wild boar Sus scrofa (L.): agricultural fields and edges. European Journal of Wildlife Research, 55(5), 517-523. doi:10.1007/s10344-009-0268-1 Crossref ● Google Scholar | ||||
| ||||
| Thurfjell, H., Spong, G., & Ericsson, G. (2013). Effects of hunting on wild boar Sus scrofa behaviour. Wildlife Biology, 19(1), 87-93. doi:10.2981/12-027 Crossref ● Google Scholar | ||||
| ||||
| Tixier, H., Duncan, P., Scehovic, J., Yant, A., Gleizes, M., & Lila, M. (1997). Food selection by European roe deer (Capreolus capreolus): effects of plant chemistry, and consequences for the nutritional value of their diets. Journal of Zoology, 242(2), 229-245. doi:10.1111/j.1469-7998.1997.tb05799.x Crossref ● Google Scholar | ||||
| ||||
| Touzot, L., Schermer, É., Venner, S., Delzon, S., Rousset, C., Baubet, É., Gaillard, J., & Gamelon, M. (2020). How does increasing mast seeding frequency affect population dynamics of seed consumers? Wild boar as a case study. Ecological Applications, 30(6), e02134. doi:10.1002/eap.2134 Crossref ● PubMed ● Google Scholar | ||||
| ||||
| Wall, W., Belisle, M., & Luke, L. A. (2010). Moose (Alces alces) wildlife habitat decision aid. Journal of Ecosystems and Management, 11(3), 45-49. doi:10.22230/jem.2010v11n3a46 Crossref ● Google Scholar | ||||
| ||||
| Zagorodniuk, I., Gavrilyuk, M., Drebet, M., Skilsky, I., Andrusenko, A., & Pirkhal, A. (2014). Wildcat (Felis silvestris schreber, 1777) in Ukraine: modern state of the population and eastwards expansion of the species. Studia Biologica, 8(3-4), 233-254. doi:10.30970/sbi.0803.372 Crossref ● Google Scholar | ||||
| ||||
| Zahorodniuk, I., & Yemelianov, I. (2012). Taxonomy and nomenclature of mammals of Ukraine. Proceedings of the National Museum of Natural History, 10, 5-30. (In Ukraine) Google Scholar | ||||
| ||||
| Zwerts, J. A., Stephenson, P. J., Maisels, F., Rowcliffe, M., Astaras, C., Jansen, P. A., ... & van Kuijk, M. (2021). Methods for wildlife monitoring in tropical forests: comparing human observations, camera traps, and passive acoustic sensors. Conservation Science and Practice, 3(12), e568. doi:10.1111/csp2.568 Crossref ● Google Scholar | ||||
Refbacks
- There are currently no refbacks.
Copyright (c) 2026 Yurii Strus, Vasylyna Strus, Ihor Dykyy, Antonina Tarasova

This work is licensed under a Creative Commons Attribution 4.0 International License.
